In addition to the enzymes mentioned earlier, the ligninolytic system of white rot fungi includes extracellular H2O2 generating enzymes, essential for peroxidase activity. To date, a number of oxidase enzymes have been shown to lead to the production of H2O2, including glyoxal oxidase (GLOX), glucose 1-oxidase, methanol oxidase, and aryl-alcohol oxidase (AO) (Zhao and Janse 1996). White rot fungi that lack H2O2 generating oxidases may rely on the oxidation of physiological organic acids such as oxalate and glyoxalate, which indirectly results in H2O2 production (Urzua et al. 1995).
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